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What makes populations stabilize? What makes them fluctuate? Are populations in complex ecosystems more stable than populations in simple ecosystems?. Titre/Auteur(s), Stability and complexity in model ecosystems [Livre] / by Robert M . May. Adresse bibliographique, Princeton: Princeton University Press, No complexity–stability relationship in empirical ecosystems (a) Equivalence between Ecopath and Lotka–Volterra models: simplified.In this case, stability increases with d, regardless of the value of n e and it does so additively (at least in cases where d is very small) in comparison with the criterion for isolated ecosystems. In this case, environmental heterogeneity is not important—emigration alone is the factor improving the stability of meta-ecosystems and its importance will depend on its magnitude relative to Cited by: Stability and complexity in model meta-ecosystems Dominique Gravel 1, *, Franc ¸ ois Massol 2,3, * & Mathew A. Leibold 4 The diversity of life and its organization in networks of interacting. that stability results from the complexity of natural ecosystems. May challenged this view in the ’sc by showing mathematically that in model ecosystems stability decreases with the number of species in a community and can therefore not be regarded as a straightforward consequence of complexity.